1 Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Canada
2 Department of Molecular Genetics, University of Toronto, Toronto, ON, Canada
3 Department of Molecular and Cellular Physiology, Systems Biology and Physiology Program, University of Cincinnati, Cincinnati, OH, USA
4 Program in Systems Biology, University of Massachusetts Medical School, Worcester, MA, USA
5 Center for Autoimmune Genomics and Etiology (CAGE) and Divisions of Biomedical Informatics and Developmental Biology, Cincinnati Children’s Hospital Medical Center, Department of Pediatrics, University of Cincinnati, Cincinnati, OH, USA
6 Canadian Institutes For Advanced Research, Toronto, Canada
*these authors contributed equally
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